[82] In the Chinese classic novel Journey to the West (16th century), the protagonist Priest of Tang was named the Golden Cicada. – Ziarat", "Ogweñ•yó'da' déñ'se' Hanadagá•yas: The Cicada and George Washington", "Great Damage from Insects; They Are Doing More Destruction to Crops This Year than Usual", "Synchronization of periodical cicada emergences", Song recordings and information of cicadas of the United States and Canada, Greater Cincinnati Cicada Information & Teaching Resources, DrMetcalf: a resource on cicadas, leafhoppers, planthoppers, spittlebugs, and treehoppers, https://en.wikipedia.org/w/index.php?title=Cicada&oldid=1014364860, Articles containing Spanish-language text, Articles containing simplified Chinese-language text, Articles containing traditional Chinese-language text, Creative Commons Attribution-ShareAlike License. The Upper Permian Dunstaniidae are found in Australia and South Africa, and also in younger rocks from China. Some species have much longer lifecycles, such as the North American genus, Magicicada, which has a number of distinct "broods" that go through either a 17-year, or in some parts of the region, a 13-year lifecycle. The cicada is also mentioned in Hesiod's Shield (ll.393–394), its voice sings when millet first ripens. In the Chinese species Subpsaltria yangi both males and female can stridulate. Fossil Cicadomorpha first appeared in the Late Triassic. Other assassin bugs include the kissing bug and wheel bug. Their dorsum is convex, lightly colored without cross striations. Versatile aggressive mimicry of cicadas by an Australian predatory katydid. They typically live in trees, feeding on watery sap from xylem tissue and laying their eggs in a slit in the bark. They also have three small ocelli located on the top of the head in a triangle between the two large eyes; this distinguishes cicadas from other members of the Hemiptera. In contrast, some small species have songs so high in pitch that they are inaudible to humans. This is the SpellCHEX dictionary for online spell checking. Another well-known song, "La Cigarra" ("The Cicada"), written by Raymundo Perez Soto, is a song in the mariachi tradition that romanticises the insect as a creature that sings until it dies. Cicadas are a frequent subject of haiku, where, depending on type, they can indicate spring, summer, or autumn. [12] About 100 species occur in the Palaearctic. Another defining characteristic is the adaptations of the fore limbs of nymphs for underground life. These periodical cicadas have an extremely long lifecycle of 13 or 17 years, with adults suddenly and briefly emerging in large numbers. Notonectidae is a cosmopolitan family of aquatic insects in the order Hemiptera, commonly called backswimmers because they swim upside down. [14] The tettigarctid (or hairy) cicadas Tettigarcta crinita of Australia and T. tomentosa have rudimentary tymbals in both sexes and do not produce airborne sounds. The short antennae protrude between the eyes or in front of them. [61], Cicadas use a variety of strategies to evade predators. The pitch is nearly constant, the sound is continuous to the human ear, and cicadas sing in scattered groups. Moulds, M.S. [56] They have the usual insect modes of locomotion, walking and flight. [37] Kanakia gigas and Froggattoides typicus are among the few that are known to be truly nocturnal and there may be other nocturnal species living in tropical forests. Cicadas are eaten by humans in various countries, including China, where the nymphs are served deep-fried in Shandong cuisine. Notonectidae is a cosmopolitan family of aquatic insects in the order Hemiptera, commonly called backswimmers because they swim upside down. As well as being coloured like tree bark, they are disruptively patterned to break up their outlines;[64] their partly transparent wings are held over the body and pressed close to the substrate. More than forty species from five genera populate New Zealand, ranging from sea level to mountain tops, and all are endemic to New Zealand and its surrounding islands (the Kermadec Islands, the Chatham Islands). [59], Several fungal diseases infect and kill adult cicadas, while other entomopathogenic fungus, Ophiocordyceps and Isaria spp., attacks nymphs. Such a rapid loss of water can be sustained only by feeding on water-rich xylem sap. The long lifecycles may have developed as a response to predators, such as the cicada killer wasp and praying mantis. This lets us find the most appropriate writer for any type of assignment. The unusual duration and timing of their emergence may reduce the number of cicadas lost to predation, both by making them a less reliably available prey (so that any predator who evolved to depend on cicadas for sustenance might starve waiting for their emergence), and by emerging in such huge numbers that they will satiate any remaining predators before losing enough of their number to threaten their survival as a species. It might seem impossible to you that all custom-written essays, research papers, speeches, book reviews, and other custom task completed by our writers are both of high quality and cheap. Cicadas have been featured in literature since the time of Homer's Iliad, and as motifs in art from the Chinese Shang dynasty. When their temperature rises above about 39 °C, they suck excess sap from the food plants and extrude the excess water through pores in the tergum at a modest cost in energy. [69], Predators such as the sarcophagid fly Emblemasoma hunt cicadas by sound, being attracted to their songs. [78] In France, the cicada represents the folklore of Provence and the Mediterranean cities. [19], The Palaeontinidae or "giant cicadas" come from the Jurassic and Lower Cretaceous of Eurasia and South America. [98][99], Temporal range: {{Fossil This allows for regulation of the size of the air bubbles and their concentration of oxygen.[4][5]. [88], In Kapampangan mythology in the Philippines, the goddess of dusk, Sisilim, is said to be greeted by the sounds and appearances of cicadas whenever she appears. [79], The cicada has represented insouciance since classical antiquity. 4. The family Cicadidae is subdivided into the subfamilies Cicadinae, Tibicininae (or Tettigadinae), Tettigomyiinae, and Cicadettinae;[4][5] they are found on all continents except Antarctica. In times of mass emergence of cicadas, various amphibians, fish, reptiles, mammals, and birds change their foraging habits so as to benefit from the glut. The Australian greengrocer, Cyclochila australasiae, is among the loudest insects in the world.[16]. However, they do not walk or run well, and take to the wing to travel distances greater than a few centimetres. The best-known genus of backswimmers is Notonecta – streamlined, deep-bodied bugs up to 1.6 cm (0.6 in) long, green, brown, or yellowish in colour. At least 3000 cicada species are distributed worldwide with the majority being in the tropics. [45], For the human ear, telling precisely where a cicada song originates is often difficult. Most genera are restricted to a single biogeographical region and many species have a very limited range. The adaptive significance is unclear as the calls are not amplified or modified by the burrow structure but it is thought that this may avoid predation. [36] Most cicadas are diurnal in their calling and depend on external heat to warm them up, while a few are capable of raising their temperatures using muscle action and some species are known to call at dusk. [73] Shaun Tan's illustrated book Cicada tells the story of a hardworking but underappreciated cicada working in an office. The wing venation varies between species and may help in identification. The abdomen is segmented, with the hindermost segments housing the reproductive organs, and terminates in females with a large, saw-edged ovipositor. Nymphs have strong front legs for digging and excavating chambers in close proximity to roots where they feed on xylem sap. The thread-legged bug is a kind of assassin bug. 1. [27] Bacteria landing on the wing surface are not repelled; rather, their membranes are torn apart by the nanoscale-sized spikes, making the wing surface the first-known biomaterial that can kill bacteria. The superfamily Palaeontinoidea contains three families. [48], Most cicadas go through a lifecycle that lasts from two to five years. The relict family Tettigarctidae differs from the Cicadidae in having the prothorax extending as far as the scutellum, and by lacking the tympanal apparatus. Elliott, Lang, and Wil Hershberger. [62] Many are extremely well camouflaged[62][63] to evade predators such as birds that hunt by sight. [25], The surface of the fore wing is superhydrophobic; it is covered with minute, waxy cones, blunt spikes that create a water-repellent film. Both males and females produce vibrations that are transmitted through the tree substrate. In addition to the mating song, many species have a distinct distress call, usually a broken and erratic sound emitted by the insect when seized or panicked. During sound production, the temperature of the tymbal muscles was found to be significantly higher. [44] The song is loud enough to cause permanent hearing loss in humans should the cicada be at "close range". Males also produce encounter calls, whether in courtship or to maintain personal space within choruses. Where multiple species occur, the species may use different heights and timing of calling. Their front tarsi are not scoop-shaped and their hind legs are fringed for swimming. In the process, their bodies and interior of the burrow become coated in anal fluids. in Burmese amber, Dominicicada youngi n. gen., n. sp. "How Do Cicadas Know When to Emerge from the Ground? In males, the abdomen is largely hollow and used as a resonating chamber. [91] Female cicadas are prized for being meatier. In other species, the males move from place to place, usually with quieter calls while searching for females. WORDS.TXT - Free ebook download as Text File (.txt), PDF File (.pdf) or read book online for free. [25] Although in general, the feeding activities of the nymphs do little damage, during the year before an outbreak of periodic cicadas, the large nymphs feed heavily and plant growth may suffer. [29][30] Some non-desert cicada species such as Magicicada tredecem also cool themselves evaporatively, but less dramatically. The middle thoracic segment has an operculum on the underside, which may extend posteriorly and obscure parts of the abdomen. Argentina. Cicadas have prominent eyes set wide apart, short antennae, and membranous front wings. [83], In Japan, the cicada is associated with the summer season. [9] There are several hundred described species in Australia and New Zealand,[c] around 150 in South Africa, over 170 in America north of Mexico,[10] at least 800 in Latin America,[11] and over 200 in Southeast Asia and the Western Pacific. Some cicadas however, have mechanisms for stridulation, sometimes in addition to the tymbals. Some cicada species play dead when threatened. [75], Cicadas are featured in the well-known protest song "Como La Cigarra" ("Like the Cicada") written by Argentinian poet and composer María Elena Walsh. The vast majority of species are active during the day as adults, with some calling at dawn or dusk. [CHEX %PARSER=2.13 %FLOATED=19991204 %GENERATED=DR/ALL %BOUND=TRUE] [2][3][b], The superfamily Cicadoidea is a sister of the Cercopoidea (the froghoppers). [80], The cicada symbolises rebirth and immortality in Chinese tradition. One genus, the periodical cicadas spend most of their lives as underground nymphs, emerging only after 13 or 17 years. lyricen var. They can inflict a painful "bite" on a human being, actually a stab with their sharp tubular mouthparts (proboscis). [87] Eventually, he became so tiny and shriveled that he turned into the first cicada. The earliest known fossil Cicadomorpha appeared in the Upper Permian period; extant species occur all around the world in temperate to tropical climates. [28], Desert cicadas such as Diceroprocta apache are unusual among insects in controlling their temperature by evaporative cooling, analogous to sweating in mammals. In the final nymphal instar, they construct an exit tunnel to the surface and emerge. [25] The postclypeus is a large, nose-like structure that lies between the eyes and makes up most of the front of the head; it contains the pumping musculature. When the droplets coalesce, they leap several millimetres into the air, which also serves to clean the wings. Assassin Bug, Family Reduviidae. By evaporative cooling, desert cicadas can reduce their bodily temperature by some 5 °C. [68] The majority of cicadas are diurnal and rely on camouflage when at rest, but some species use aposematism-related Batesian mimicry, wearing the bright colors that warn of toxicity in other animals; the Malaysian Huechys sanguinea has conspicuous red and black warning coloration, is diurnal, and boldly flies about in full view of possible predators. [60] This fungus is also capable of dosing cicadas with psilocibin, the psychedelic drug found in magic mushrooms, as well as cathinone, an alkaloid similar to various amphetamines. A few species are found in southern Europe,[7] and a single species was known from England, the New Forest cicada, Cicadetta montana, which also occurs in continental Europe. This high degree of endemism has been used to study the biogeography of complex island groups such as in Indonesia and Asia. The cicada also modulates the song by positioning its abdomen toward or away from the substrate. [92] The 17-year "Onondaga Brood"[93] Magicicada is culturally important and a particular delicacy to the Onondaga people. A phylogenetic treatment suggested by a 2018 study. [96], Cicadas are not major agricultural pests, but in some outbreak years, trees may be overwhelmed by the sheer numbers of females laying their eggs in the shoots. The superfamily is divided into two families, Tettigarctidae, with two species in Australia, and Cicadidae, with more than 3,000 species described from around the world; many species remain undescribed. [87] The Greeks also used a cicada sitting on a harp as an emblem of music. [18] However, this group is currently thought to be more distantly related to Cicadomorpha than previously thought. The song was famously recorded by Mercedes Sosa, among other Latin American musicians. They are all predators and typically range from 0.5 to 1.5 cm (0.2–0.6 in) in length. Zootaxa 4438(3): 443–470. [3] Another of the same region, N. maculata, is distinguished by its mottled brick-coloured forewings. Cheap paper writing service provides high-quality essays for affordable prices. 2007. A A's AMD AMD's AOL AOL's AWS AWS's Aachen Aachen's Aaliyah Aaliyah's Aaron Aaron's Abbas Abbas's Abbasid Abbasid's Abbott Abbott's Abby Abby's Abdul Abdul's Abe Abe's Abel Abel's BBC Science and nature factfiles: Water boatman, common backswimmer, physorg.com: Gold Strike in Science World, Photo of Anisops assimilis swimming by Paddy Ryan, BBC Radio special on the sounds made by water boatmen, https://en.wikipedia.org/w/index.php?title=Notonectidae&oldid=1009234537, Short description is different from Wikidata, Articles with dead external links from February 2021, Creative Commons Attribution-ShareAlike License, This page was last edited on 27 February 2021, at 13:30. [67] Since the cicadas in any given area exceeds the number predators can eat, all available predators are sated, and the remaining cicadas can breed in peace. range|Late Triassic|Recent}}, The Auchenorrhyncha were formerly part of the obsolete ". [23] The male abdomen in some species is largely hollow, and acts as a sound box. 2009. [45] Shells of cicadas are employed in traditional Chinese medicines. A further 300 collected Australian species remain to be described. [81] In the Chinese essay "Thirty-Six Stratagems", the phrase "to shed the golden cicada skin" (simplified Chinese: 金蝉脱壳; traditional Chinese: 金蟬脫殼; pinyin: jīnchán tuōqiào) is the poetic name for using a decoy (leaving the exuvia) to fool enemies. [23] The Auchenorrhyncha differ from other hemipterans by having a rostrum that arises from the posteroventral part of the head, complex sound-producing membranes, and a mechanism for linking the wings that involves a down-rolled edging on the rear of the fore wing and an upwardly protruding flap on the hindwing. Though these cicadas have lifecycles that can vary from one to nine or more years as underground larvae, their emergence above ground as adults is not synchronized, so some members of each species appear every year.[1]. [95] Male cicadas produce very loud calls that can damage human hearing. These chemicals alter the behaviour of the cicadas, driving males to copulate, even with males, and is thought to be beneficial to the fungus as the fungal spores are dispersed by a larger number of infected carriers. 2018. [90] Cicadas are also eaten in Malaysia, Burma, North America, and central Africa, as well as the Balochistan region of Pakistan, especially in Ziarat. There are about 350 species in two subfamilies: Notonectinae with seven genera, and Anisopinae with four genera. lyricen")", "The Ecology, Behavior, And Evolution of Periodical Cicadas", "Exploitation of Sexual Signals by Predators and Parasitoids", "La Fontaine, fabuleusement inspiré par Esope – Un autre regard sur la Grèce", "Cicada in Chinese Folklore (with bibliography)", "The Cicadas' Song: Japan's Summer Soundtrack", "Formation of the World, Kapampangan Mythology", "Taazan – Cicada – How to Cook ? Branches of young trees may die as a result. By rapidly vibrating these membranes, a cicada combines the clicks into apparently continuous notes, and enlarged chambers derived from the tracheae serve as resonance chambers with which it amplifies the sound. [9], Cicadas are commonly eaten by birds and sometimes by squirrels,[57] as well as bats, wasps, mantises, spiders, and robber flies. [85] According to Lafcadio Hearn, the song of Meimuna opalifera, called tsuku-tsuku boshi, is said to indicate the end of summer, and it is called so because of its particular call. Members in the former subfamily are often larger than those in the latter.[2]. In wet habitats, larger species construct mud towers above ground to aerate their burrows. [18] The first of these was a fore wing discovered in the Taynton Limestone Formation of Oxfordshire, England; it was initially described as a butterfly in 1873, before being recognised as a cicada like form and renamed Palaeontina oolitica. This species is usually about 1 to 1.5 inches in length, but is so thin that it appears to be no more than a slender stick. One species is found on Norfolk Island which technically is part of Australia. [31] Conversely, many other cicadas can voluntarily raise their body temperatures as much as 22 °C (40 °F) above ambient temperature. [53], An adult cicada that has just emerged and is waiting to dry before flying away, Newly emerged adult cicada held on human fingers, Cicada exuviae after the adult cicada has left, Cicada nymphs drink sap from the xylem of various species of trees, including oak, cypress, willow, ash, and maple. [34], Average temperature of the natural habitat for the South American species Fidicina rana is about 29 °C (84 °F). lakes, ponds, marshes, and are sometimes found in garden ponds and even swimming pools. Many of them have common names such as cherry nose, brown baker, red eye, greengrocer, yellow Monday, whisky drinker, double drummer, and black prince. [70] Singing males soften their song so that the attention of the listener gets distracted to neighbouring louder singers, or cease singing altogether as a predator approaches. Most cicadas are cryptic. [54][55], Cicadas, unlike other Auchenorrhyncha, are not adapted for jumping (saltation). Some species also have courtship songs, generally quieter, and produced after a female has been drawn to the calling song. In the absence of rain, dew condenses on the wings. 4th Salon "RHODOS" - Acceptances Last Update March 17 2021. Rain rolls across the surface, removing dirt in the process. Some previous works also included a family-level taxon called the Tibiceninae. [26], The thorax has three segments and houses the powerful wing muscles. [35] Many cicadas sing most actively during the hottest hours of a summer day; roughly a 24-hour cycle. They have also been used in myth and folklore as symbols of carefree living and immortality. [33] The sounds may further be modulated by membranous coverings and by resonant cavities. Large cicadas can fly rapidly to escape if disturbed. Assassin bugs are the death-dealing ninjas of the insect world, and their camouflage and stealth are legendary. Cicadas live underground as nymphs for most of their lives at depths down to about 2.5 m (8 ft). Cicada fossils (Cicadoidea: Tettigarctidae and Cicadidae) with a review of the named fossilised Cicadidae. The mouthparts form a long, sharp rostrum that they insert into the plant to feed. [74] Branden Jacobs-Jenkins' play Appropriate takes place on an Arkansas farm in summer, and calls for the sounds of mating cicada's to underscore the entire show. [d] They are described by Aristotle in his History of Animals and by Pliny the Elder in his Natural History; their mechanism of sound production is mentioned by Hesiod in his poem "Works and Days": "when the Skolymus flowers, and the tuneful Tettix sitting on his tree in the weary summer season pours forth from under his wings his shrill song". Only a rare few species are known to be nocturnal. [77], Cicadas have been used as money, in folk medicine, to forecast the weather, to provide song (in China), and in folklore and myths around the world. [67], Cicadas have been featured in literature since the time of Homer's Iliad, and as motifs in decorative art from the Chinese Shang dynasty (1766–1122 BC.). [52] An alternate hypothesis is that these long lifecycles evolved during the ice ages so as to overcome cold spells and that as species co-emerged and hybridized they left distinct species that did not hybridize having periods matching prime numbers. They are considered as representing the original state from which other cicada communication has evolved. In contrast to other aquatic insects that cling to submerged objects, Anisops deanei uses a unique system to stay submerged: using the extra oxygen supply from haemoglobin in their abdomen, instead of using oxygen dissolved in the water. Males disable their own tympana while calling, thereby preventing damage to their hearing;[43] a necessity partly because some cicadas produce sounds up to 120 dB (SPL)[43] which is among the loudest of all insect-produced sounds. They are in the suborder Auchenorrhyncha,[a] along with smaller jumping bugs such as leafhoppers and froghoppers. [9] After mating, the female cuts slits into the bark of a twig where she deposits her eggs. [42], Although only males produce the cicadas' distinctive sounds, both sexes have membranous structures called tympana by which they detect sounds, the equivalent of having ears. PLoS One 4(1): e4185. The Tettigarctidae differ from other cicadas in producing vibrations in the substrate rather than audible sounds. Many of the North American species are the annual or jarfly or dog-day cicadas, members of the Neotibicen, Megatibicen, or Hadoa genera, so named because they emerge in late July and August. This page was last edited on 26 March 2021, at 17:41. Milkweed assassin bugs are out for blood whether it be that of other insects or even humans. They have an exceptionally loud song, produced in most species by the rapid buckling and unbuckling of drumlike tymbals. [13] Many species await formal description and many well-known species are yet to be studied carefully using modern acoustic analysis tools that allow their songs to be characterized. Historical Biology, 24, 461–466. [7][24] Cicadas have prominent compound eyes set wide apart on the sides of the head. Jean de La Fontaine began his collection of fables Les fables de La Fontaine with the story "La Cigale et la Fourmi" ("The Cicada and the Ant") based on one of Aesop's fables; in it, the cicada spends the summer singing, while the ant stores away food, and finds herself without food when the weather turns bitter. [76], The Brazilian artist Lenine with his track "Malvadeza" from the album "Chão" creates a song built upon the sound of the cicada that can be heard along the track. [6] Cicadas are also notable for the great length of time some species take to mature.[7]. The name is directly from the onomatopoeic Latin cicada. A loud cicada song, especially in chorus, has been asserted to repel predators, but observations of predator responses refute the claim. Professional academic writers. The Upper Triassic Mesogereonidae are found in Australia and South Africa. One fossil genus and species (Burmacicada protera) based on a first-instar nymph has recently been reported from 98–99M years ago in the Late Cretaceous,[22] although questions remain about its assignment to the Cicadidae. [14] The best-known North American genus, however, may be Magicicada. Introducing cicadas – Cicadas – Te Ara Encyclopedia of New Zealand, "Jurassic Palaeontinidae From China and the Higher Systematics of Palaeontinoidea (Insecta: Hemiptera: Cicadomorpha)", https://doi.org/10.11646/zootaxa.4438.3.2, https://doi.org/10.1080/08912963.2011.603421, "Researchers find cicada wing structure able to kill bacteria on contact", "Evaporative cooling in the desert cicada: thermal efficiency and water/metabolic costs", "Substrate-transmitted acoustic signals of the primitive cicada, Tettigarcta crinita Distant (Hemiptera Cicadoidea, Tettigarctidae)", "Mechanical properties of glycerinated fibres from the tymbal muscles of a Brazilian cicada", "Sound of Insect Music Ushers in Fall Season", "Cicada acoustic communication: potential sound partitioning in a multispecies community from Mexico (Hemiptera: Cicadomorpha: Cicadidae)", "No acoustic benefit to subterranean calling in the cicada, "Global Cicada Sound Collection I: Recordings from South Africa and Malawi by B. W. Price & M. H. Villet and harvesting of BioAcoustica data by GBIF", "Sequence A161664, Safe periods for the emergence of cicada species on prime number cycles", "Scanjet image of 7-petal flower from Jade plant approximately 25 years or more years old in Livermore, California", "Jumping mechanisms of treehopper insects (Hemiptera, Auchenorrhyncha, Membracidae)", "Psychoactive plant- and mushroom-associated alkaloids from two behavior modifying cicada pathogens", "1-Trick Chameleon: Predators Learn to See Through Camouflage", "Subspecies Neotibicen lyricen lyricen - Lyric Cicada ("ssp. Partly by the pattern in which it combines the clicks, each species produces its own distinctive mating songs and acoustic signals, ensuring that the song attracts only appropriate mates. The common backswimmer, N. glauca, is widespread in Europe, including the United Kingdom where it is known as the greater water boatman. VILLALBA RAUL Pain, Accepted (open-monochrome)Clowns, Accepted (open-color)The escape, Accepted (open-color)The grand cross, GPU Bronze Medal (open-monochrome) Memories, Accepted (open-color)The sphinx, Accepted (open-monochrome)Australia [20], Most fossil Cicadidae are known from the Cenozoic,[21] and the oldest unambiguously identified specimen is Davispia bearcreekensis (subfamily Tibicininae) from 59–56 Ma. Assassin or ambush bug identification is pretty easy -- it's finding them that's difficult. [49][50][51] A specialist predator with a shorter life cycle of at least two years could not reliably prey upon the cicadas. ", "An appraisal of the higher classification of cicadas (Hemiptera: Cicadoidea) with special reference to the Australian fauna", "A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily classification", "Biogeography of the Cicadas (Hemiptera: Cicadidae) of North America, North of Mexico", "The Current Status of Cicada Taxonomy on a Region-by-Region Basis", 1. The thread-legged assassin bug is often mistaken for a stick insect. [72] In the classic 14th-century Chinese novel Romance of the Three Kingdoms, Diaochan took her name from the sable (diāo) tails and jade decorations in the shape of cicadas (chán), which adorned the hats of high-level officials. [65][66], The periodical cicadas (Magicicada) make use of predator satiation: they emerge, all at once, at long intervals of 13 or 17 years; their juveniles are probably the longest-lived of all insect development stages. [1] They are similar in appearance to Corixidae (water boatmen), but can be separated by differences in their dorsal-ventral coloration, front legs, and predatory behavior.
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